Rune of Manos/1.03

Dark Elf
Allows for the production of Sorcerors and Spider Priestesses at the Tree of Pain.

High Elf
Allows for the production of Mystics at the Tree of Gold. If both Rune of Manos and Rune of Cielos have been researched it allows Ice Maidens from the Tree of Gold.

Wood Elf
Allows for the production of Druids at the Tree of Life.